LPR print, can it be directly to printer?

I have a USB printer, connected to a network print server (TL-PS310U). I want to print from Debian/Ubuntu to this printer. The queue is named 2409ee3c, but when I try to use lpr, it says that I need to add /version=1.1: user@user:~$ lpr -H 192.168.100.1 -P prt0 C0003022.che.spl lpr: Error - add '/version=1.1' to server name. Then I added the printer in CUPS (http://localhost:631) as lpd, with name Printserver: lpd://192.168.2.30/2409ee3c And I now can print with LPR: user@user:/run/shm$ export CUPS_SERVER=localhost:631; lpr -H 192.168.2.30 -P 2409ee3c C0003022.che.spl lpr: The printer or class does not exist. jsivil@jsivil:/run/shm$ export CUPS_SERVER=localhost:631; lpr -H 127.0.0.1 -P Printserver C0003022.che.spl My questions are: 1) Can I print with LPR directly to this printer behind a printserver? (I've tried netcat but it doesn't work for this printserver) 2) Does LPR need CUPS? Is it OK to use 127.0.0.1? 3) How can I avoid having to export the CUPS_SERVER variable prior to use lpr? Thanks