Unable to transfer files (put) using ftp or filezilla from windows to ubuntu VM

I've setup a ubuntu 64 bit VM (16.04) using Oracle VirtualBox. Using ftp, I am able to connect to the VM. I am also able to list (ls) the contents of the folder. However I am unable to put files to the VM. C:\>ftp x.x.x.x Connected to x.x.x.x. 220 (vsFTPd 3.0.3) 200 Always in UTF8 mode. User (x.x.x.x:(none)): user 331 Please specify the password. Password: 230 Login successful. ftp> quote pasv 227 Entering Passive Mode (...) ftp> put trnsfr.txt 200 PORT command successful. Consider using PASV. 550 Permission denied. On ubuntu, I set permissions on folder to rwx on owner,group,all. I then modified /etc/vsftpd.conf as follows: pasv_enable=YES pasv_min_port=30000 pasv_max_port=30100 port_enable=yes pasv_address=x.x.x.x allowed data connections and restarted vsftpd daemon: iptables -I INPUT -p tcp --destination-address 30000:30100 -j ACCEPT /etc/init.d/vsftpd restart but still permission error exists. I then tried using filezilla: The connections is successful but again files can not be transferred: with dft settings in vsftpd: Response: 550 Permission denied. Error: Critical file transfer error with the settings in vsftpd.conf as above: the following is displayed in filezilla: Error: The data connection could not be established: WSAEADDRNOTAVAIL - Cannot assign requested address I believe the problem is to do with the data connection - but am not sure what/how to resolve?